The relation of language variation to reconstructed languages and to the methodology of reconstruction has long been neglected. The articles in the present volume consider this relationship from a number of different angles, with a number of different focuses. Several of the papers discuss evidence from Germanic, either Proto-Germanic (Joseph, Schwink), or daughter languages such as Dutch (Goss & Howell), Afrikaans (Roberge), Newcastle English (Milroy), and a Wisconsin German dialect (Geiger & Salmons). Other papers look at Italian (Cravens), Spanish (Harris-Northall), and the non-Indo-European languages or families Aramaic (Miller), and Proto-Hmong-Mien (Ratliff), and the Southeast Asian languages Phan Rang Cham and Tsat (Thurgood). In doing so they bring together a number of interconnected issues which are of current concern in comparative and historical linguistics.